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Treadmill inclination and VO2Max on my FR255

I am calibrating the distance I jog on the treadmill after each session.

Historically, I have jogged with a 3% inclination. Can I confirm the following:

1) everything else equal, Garmin's algo is probably underestimating my VO2Max.

2) I am planning to reduce the inclination to 0. Everything else equal, the same jog will now be done at a lower HR, and this should have the effect to adjust Garmin's estimate of my VO2Max to a higher, more realistic level.

Is this correct ?

The reason for my question: VO2Max is given in ml/min/kg (or in plain English, in milliliter of O2 metabolised per minute per kg of weight). In my jog, what is the variable that allows this ml/min/kg number to go up when my HR goes down ? I need to clarify this connection.